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Has he had full blood work done besides just the tick tests?
Has he had xrays to evaluate his joints and lungs? Often difficult diseases or autoimmune conditions can present like your dogs symptom and if so we need to evaluate his skeletal system.
It is also not wrong to consider getting an internal medicine specialist involved. This case is not going to be an easy one to diagnose it sounds like. With an internal medicine specialist involved we may be able to get to the bottom of things much faster.
Can you send me your dog's labwork and tests that have been run so i can take a look?
ok I will keep an eye on this post for the results