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You have all that the cylinders need to run fuel and spark and compression. Can you check for a valve train issue that is not allowing the valves to open on these cylinders. You can do that with a visual inspection or by applying compressed air to the cylinders and cranking the engine by hand slow to see if you get air flow out the exhaust and throttle body based on the stroke its on.
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I am not saying it is a compression issue. I am saying it looks like the valves are not opening on these cylinders possibly from worn cam lobes for example.