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The scope of my question is limited to inheritance taxes, not estate taxes.
I assume you are not a resident of PA.
Another expert here:
Since there are no assets that are being probated, the PA inheritance taxes could not be paid by the estate. The inheritance tax on the POD CDs would have to be paid by the recipients of those assets. You as trustee would be responsible for paying the inheritance tax due on the trust assets from the funds in the trust.