Many times, things like this are caused from food allergies. Cats are often allergic to corn or corn-gluten which are regularly used in commercial foods. I would try putting him on a corn-free diet such as Wellness, Innova, Felidae or California Natural. Make the change gradually, by adding a little new food to his existing diet. Each day add a little more new food and a little less of the old food until he is eating all new food in about a week to 10 days. Once he's on the new food for a couple of weeks, you should (hopefully) see a big improvement in the problem's he's having with his feet.
Please let me know how he does and if I can help further.